I have little experience with the RNS315 other than playing around with it in the show room while waiting for service. But, the little time I spent with it turned me off. It has a small screen, weird controlls, low resolution, and limited functionality.
When I purchased my old 06 Jetta TDI Pkg 3, it has the old nav. After about two years dealing with that, I bought the RNS-510 and swapped them out. It was a good choice for me. It has better resoultion, larger screen, hard drive for music, better bluetooth functionality (on screen phone books and dial pad), can play DVD movies, and just performs better.
It should be mentioned that my car is like my office, and I have appointments throughout the day, sometimes 30-50 miles apart. So, an in dash nav is the best solution for me. It might not be for you. If you will only be casually using your nav, you're better off with a portable nav system that you can move from car to car and not have in the car when you don't need it (and save yourself $1,000+).
I was able to update the software / firmware on my older RNS 510 and now have the speed limits desplayed on screen and all the other newer features. When we bought my wife's Tiguan, they had two in our color choice: One with the big roof that looks like it will definately break, or, one without the roof, but with the RNS-510 (with back-up camera). We took the one with the RNS-510 and she loves it (along with the iPod functionality).